The milestone vehicle was a Freelander 2 TD4_e in Biscay Blue and Land Rover says it will sell the model in the UK market.
The Halewood plant has been producing the Freelander 2 for three years since production was moved from Birmingham when the new model was introduced.
Production at the Liverpool plant has increased steadily since 2007 and is expected to increase by around 31 per cent in 2010.
This has been encouraged by strong sales for the 4×4 manufacturer. In April, Land Rover recorded the biggest increase in sales in its 62-year history.
Land Rover says impressive sales meant production of the Freelander 2 at the Halewood plant was increased by 20 per cent last year to meet demand.
In a further boost to the plant, which phased out production of the Jaguar X-Type at the end of last year, production of the ‘baby’ Range Rover, the LRX, will begin next year.
The production of the LRX and the continued success of the Freelander 2 has secured over 2,000 jobs at the plant.
Halewood Operations Director Michael Straughan said, “This achievement is thanks to the Halewood workforce, who have continuously driven improvements since the launch of Freelander 2 to maximise customer satisfaction.
"I’d like to congratulate the Halewood workforce on this team achievement.”
The Freelander 2 is a groundbreaking model for the manufacturer and SUV market as a whole. In 2009, it became the first SUV to fit fuel-saving StopStart technology as standard on diesel manual vehicles.
Prices for the Freelander 2 start from £21,295.